Coach Eddie Jones has named a full-strength England lineup to take on Tonga in their World Cup opener in Sapporo on Sunday.
Owen Farrell will lead the side featuring 13 of the 15 players that started against Ireland at Twickenham last month. Jones, though, has opted to start Farrell at inside centre again with George Ford retaining the No 10 jersey.
Wing Joe Cokanasiga and lock George Kruis were left out of the run-on XV and replaced by Anthony Watson and Courtney Lawes.
Joe Marler, who came of retirement in July, continues at loosehead prop in the absence of Mako Vunipola, who is still battling to overcome a hamstring injury.
The match kicks off at 12:15pm (South African time) on Sunday.
England – 15 Elliot Daly, 14 Anthony Watson, 13 Manu Tuilagi, 12 Owen Farrell (c), 11 Jonny May, 10 George Ford, 9 Ben Youngs, 8 Billy Vunipola, 7 Sam Underhill, 6 Tom Curry, 5 Maro Itoje, 4 Courtney Lawes, 3 Kyle Sinckler, 2 Jamie George, 1 Joe Marler
Subs: 16 Luke Cowan-Dickie, 17 Ellis Genge, 18 Dan Cole, 19 George Kruis, 20 Lewis Ludlam, 21 Willi Heinz, 22 Henry Slade, 23 Jonathan Joseph
Photo: England Rugby website