SA-born prop WP Nel returns as one of an experienced Scotland trio for the Six Nations clash with England at Murrayfield on Saturday.
Fullback Blair Kinghorn, recovering from a knee injury, is also back, along with Nel, a linchpin of the Scottish scrum for nearly a decade.
The veteran front-row forward makes a comeback from a neck injury to replace Javan Sebastian.
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Additionally, flanker Hamish Watson receives a recall due to a concussion injury to Josh Bayliss, while Magnus Bradbury has also been roped in by head coach Gregor Townsend.
Ross McCann exits the squad, while the injured Darcy Graham remains sidelined with a groin problem.
Full Scotland squad – Forwards: Ewan Ashman, Magnus Bradbury, Jamie Bhatti, Alex Craig, Andy Christie, Scott Cummings, Jack Dempsey, Rory Darge, Grant Gilchrist, Matt Fagerson, Zander Fagerson, Alec Hepburn, Johnny Matthews, Elliot Millar-Mills, WP Nel, Jamie Ritchie, Pierre Schoeman, Sam Skinner, George Turner, Hamish Watson, Glen Young. Backs: Ben Healy, George Horne, Rory Hutchinson, Huw Jones, Blair Kinghorn, Stafford McDowall, Harry Paterson, Ali Price, Cameron Redpath, Arron Reed, Kyle Rowe, Finn Russell, Kyle Steyn, Ross Thompson, Sione Tuipulotu, Duhan van der Merwe, Ben White.
Photo: @Scotlandteam/Twitter